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BIOLOGICAL AND MEDICAL
PHYSICS
BIOMEDICAL ENGINEERING
Springer
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BIOLOGICAL AND MEDICAL
PHYSICS
BIOMEDICAL ENGINEERING
The fields of biological and medical physics and biomedical engineering are
broad, multidisciplinary and dynamic. They lie at the crossroads of frontier
research in physics, biology, chemistry, and medicine. The Biological & Medical
Physics/Biomedical Engineering Series is intended to be comprehensive, covering
a broad range of topics important to the study of the physical, chemical and
biological sciences. Its goal is to provide scientists and engineers with textbooks,
monographs, and reference works to address the growing need for information.
Editor-in-Chief:
Elias Greenbaum, Oak Ridge National Laboratory, Oak Ridge, Tennessee, USA
Editorial Board:
Masuo Aizawa, Department of Bioengineering, Tokyo Institute of Technology,
Yokohama, Japan
Norma Allewell, Department of Biochemistry, University of Minnesota, St. Paul,
Minnesota, USA
Olaf S. Andersen, Department of Physiology, Biophysics and Molecular
Medicine, Cornell University, New York, USA
Robert H. Austin, Department of Physics, Princeton University, Princeton, New
Jersey, USA
James Barber, Department of Biochemistry, Imperial College of Science,
Technology and Medicine, London, England
Howard C. Berg, Department of Molecular and Cellular Biology, Harvard
University, Cambridge, Massachusetts, USA
Victor Bloomfield, Department of Biochemistry, University of Minnesota, St.
Paul, Minnesota, USA
Robert Callender, Department of Biochemistry, Albert Einstein College of
Medicine, Bronx, New York, USA
Britton Chance, Department of Biochemistry and Biophysics, University of
Pennsylvania, Philadelphia, Pennsylvania, USA
Steven Chu, Department of Physics, Stanford University, Stanford, California, USA
Louis J. DeFelice, Department of Pharmacology, Vanderbilt University, Nashville,
Tennessee, USA
Johann Deisenhofer, Howard Hughes Medical Institute, The University of Texas,
Dallas, Texas, USA
Continued on page 133 after Index
 
Howard C. Berg
E. coli in Motion
With 42 Figures, 1 in Full Color
13
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Howard C. Berg
Department of Molecular and Cellular Biology
Harvard University
Cambridge, MA 02138
USA
hberg@biosun.harvard.edu
Library of Congress Cataloging-in-Publication Data
Berg, Howard C., 1934–
E. coli in motion / Howard C. Berg.
p. ; cm.—(Biological and medical physics biomedical engineering)
Includes bibliographical references and index.
ISBN 0-387-00888-8 (hc. : alk. paper)
1. Escherichia coli. 2. Microorganisms–Motility. I. Title. II. Series.
[DNLM: 1. Escherichia coli—pathogenicity. QW 138.5.E8 B493e 2003]
QR82.E6B47 2003
579.3¢42—dc21
2003045491
ISBN 0-387-00888-8
Printed on acid-free paper.
© 2004 Springer-Verlag New York, Inc.
All rights reserved. This work may not be translated or copied in whole or in part without
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Printed in the United States of America.
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